Montana State University

Adam Richman

Adam Richman

Adam Richman Associate Professor, PhD from University of California, San Diego 1992

Office: 313 Plant BioScience Building
Lab: 339 Plant BioScience Building

Office Phone: 406-994-7750

Fax: 406-994-7600
Email Dr. Richman


Population and evolutionary genetics of self-recognition systems, island biogeography, evolutionary community ecology.


Population geneticist; Research in the evolutionary genetics of self-recognition; Teaching BIOL 301 (Genetics) and BIOL 501 (Evolutionary and Population Genetics).


  • Ph.D. Biology, University of California, San Diego 1991
  • B.A. Biology, University of California, San Diego 1985


  • Society for the Study of Evolution
  • American Genetics Association


  • 1998-2001, National Science Foundation,“Evolutionary Genetics of Self-Recognition Systems”, $300,000.


  • BIOL 301: Genetics (alternate years)
  • BIOL 501: Evolutionary and Population Genetics (alternate years)


Refereed Journals

Richman A.D., L. G. Herrera, and D. Nash (2001). MHC Class II Beta Sequence Diversity in the Deer Mouse (Peromyscus maniculatus): Implications for models of balancing selection. Molecular Ecology 10:2765-2773.

Lavin, M. , M.F. Wojciechowski, A.D. Richman, J. Rotella, M. J. Sanderson, and a.B. Matos (2001). Identifying Tertiary Radiations of Fabaceae in the Greater Antilles: alternatives to cladistic vicariance analysis. International Journal of Plant Sciences162:53-75.

Richman A.D., L. G. Herrera, and D. Nash (in press). Characterization of Peromyscus MHC Class II beta sequences by LA RT-PCR and DGGE. The European Journal of Immunogenetics.

Richman, A.D. 2000. Evolution of balanced genetic polymorphism. Molecular Ecology 9:1953-1964.

Richman, A.D. 2000. Evidence for a bottleneck in S-allele diversity in the Solanaceae. Annals of Botany 85:241-245.

Price, T.D., I. Lovett, E. Birmingham, H. L. Gibbs and A.D. Richman, 2000. The imprint of history on communities of North American and Asian warblers. The American Naturalist 156: 354-367.

Richman, A.D. and J. R. Kohn, 2000. Evolutionary genetics of self-incompatibility in the Solanaceae. Plant Molecular Biology 42: 169-179.

Richman, A.D. and J. R. Kohn, 1999. Self-incompatibility alleles from Physalis: Implications for historical inference from balanced polymorphisms. Proceedings of the National Academy of Science, USA. 96(1): 168-172.

Price, T, D., A. Helbig and A. D. Richman. 1997. Rapoport's Rule revisited. Evolution 51:552-561.

Richman, A.D. and J. R. Kohn. 1997. Self-incompatibility RNases from three plant families: homology or convergence? American Journal of Botany 84:912-917

Richman, A.D., M.K. Uyenoyama and J. R. Kohn. 1996. Contrasting patterns of S-allelic diversification in two self-incompatible species in the Solanaceae. Science 273: 1212-1216.

Richman, A.D. and J. R. Kohn. 1996. Learning from rejection: evolutionary ecology of gametophytic self-incompatibility. Trends in Ecology and Evolution 11:497-502.

Richman, A.D. 1996. Ecological divergence and community structure in the Old World Leaf Warblers (genus Phylloscopus). Evolution 50:2461-2470.

Richman, A.D., M.K. Uyenoyama, and J.R. Kohn. 1996. S-allele diversity in a natural population of ground cherry, Physalis crassifolia. Heredity 76:497-505.

Richman, A.D., T-h Kao, S. Schaeffer and M.K. Uyenoyama. 1995. S-allele sequence diversity in natural populations of Solanum carolinense (Horsenettle). Heredity 75:405-415.

Case, T.J., D.T. Bolger, and A.D. Richman. 1995. Reptilian extinctions: the last ten thousand years. Chapter in: Conservation biology: The theory and practice of nature conservation, preservation and management, P.L. Fiedler and S.K. Jain, editors. Chapman and Hall.

Alberts, A., A.D. Richman, D. Tran, R. Sauvajot, C. McCalvin, and D.T. Bolger. 1993. Effects of habitat fragmentation on populations of native and exotic plants in Southern California coastal scrub. Chapter in: Interface between ecology and land development in Southern California, (J.K. Keeley, ed.), Southern California Academy of Sciences.

Richman, A.D., and T.D. Price. 1992. The evolution of ecological differences in the Old World Leaf Warblers: roles of history and adaptation. Nature 355:817-821.

Richman, A.D., T.J. Case, and T.D. Schwaner. 1988. Natural and unnatural extinctions of reptiles on islands. The American Naturalist 131:611-630.



Evolutionary genetics of the MHC in Peromyscus


Undergraduate and graduate genetics